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Russia's first regiment of Avangard hypersonic missiles has been put into service, the defence ministry says.
The location was not given, although officials had earlier indicated they would be deployed in the Urals.
They have a "glide system" that affords great manoeuvrability and could make them impossible to defend against.
Defence Minister Sergei Shoigu confirmed the "Avangard hypersonic glide vehicle entered service at 10:00 Moscow time on 27 December", calling it a "landmark event".
Mr Putin said on Tuesday the Avangard system could penetrate current and future missile defence systems, adding: "Not a single country possesses hypersonic weapons, let alone continental-range hypersonic weapons. "
Mr Putin unveiled the Avangard and other weapons systems in his annual state-of-the-nation address in March 2018, likening it to a "meteorite" and a "fireball".
Mounted on top of an intercontinental ballistic missile, the Avangard can carry a Nuclear Weapon of up to two megatons. Russia's defence ministry has released video of the Avangard system, but weapons experts have expressed scepticism about its effectiveness.
In a statement, The Pentagon said it would "not characterise the Russian claims" about the Avangard's capabilities. The US has its own hypersonic missile programme, as does China, which in 2014 said it had conducted a Test flight of such as weapon.
The New START accord, which expires in February 2021, is The Last major nuclear arms control treaty between Russia and the US.
US President Donald Trump said he wanted a new nuclear pact to be signed by both Russia and China.
It is hard to determine if Russia's new Avangard hypersonic missile system really has entered service, as Moscow claims, or if this is just an advanced phase of field testing.
But President Putin's eagerness to claim bragging rights is to some extent justified. Russia looks to be ahead in the hypersonic stakes. China is also developing such systems; while the US appears to be somewhat behind.
Hypersonic missiles, as their name implies, fly very fast, at above Mach 5 - ie at least five times the speed of sound.
They can be cruise-type missiles, powered throughout their flight. Or, they can be carried aloft on board a ballistic missile from which the hypersonic "glide vehicle" separates And Then flies to its target.
Such "boost-glide" systems, as they are known (Avangard appears to be one of these), are launched like a traditional ballistic missile, but instead of following an arc high above the atmosphere, the re-entry vehicle is put on a trajectory that allows it to enter Earth's atmosphere quite quickly, before gliding, un-powered, for hundreds or thousands of kilometres.
It is not so much the speed of the hypersonic weapon alone that counts. It is its extraordinary manoeuvrability as it glides towards its target.
This poses a huge problem for existing anti-missile defence systems.
Indeed the glide vehicle's trajectory, "surfing along The Edge of the atmosphere" as one expert put it to me recently, presents any defensive system with additional problems.
Thus, if Russia's claims are true, it has developed a long-range intercontinental missile system that may well be impossible to defend against.
The Announcement that Avangard is operational heralds a new and dangerous era in the nuclear arms race.A display of the Avangard hypersonic boost-glide weapon
Some analysts might well see Russia's development programme as a long-term strategy to cope with Washington's abiding interest in anti-missile defences. The US argument that these are purely designed to counter missiles from "rogue-states" like Iran or North Korea has carried little weight in Moscow.
This all comes at a time when the whole network of arms control agreements inherited from the Cold War is collapsing.
With a whole new generation of nuclear weapons at the threshold of entering service, many believe not just that existing agreements should be bolstered, but that new treaties are needed to manage what could turn into a new nuclear arms race.